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Harriet Johnston has extensive work experience in various industries and roles. Harriet'smost recent role is as the Head of Brand and Marketing at Y TREE, a company specializing in financial life intelligence. Prior to this, they worked at Charlotte Street Partners, where they held the positions of Creative Director and Partner. Harriet also served as a Trustee at Street Soccer (Scotland) Ltd, a non-profit organization using football as a means of social change. Harriet gained experience in the charity sector as a Major Gifts Scotland officer at Shelter UK. Harriet'sprevious roles include being a Tutor at the University of Bologna and holding the position of Head of Communications and Operations at Agenfor Group. Harriet also worked as a Translator and Writer on a freelance basis. Early in their career, Harriet was a Producer for MTV Networks and FBC Media, where they produced news bulletins and generated economic stories for various media outlets.

Harriet Johnston completed their Bachelor's degree in Laws (LLB) from King's College London between 1996 and 2000. In 2013 and 2014, they pursued a Master's degree in Fundraising for Public Institutions and Non-profit Organizations from Alma Mater Studiorum – Università di Bologna. It is unclear when Harriet attended Downe House and St Georges School for Girls, but they did not obtain any degrees or specify any field of study during those times.