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Holly Mostoller has a diverse work experience, starting with their role as Administrative Director at Jacksonville Orthopaedic Institute from August 2004 to September 2005. Holly then moved on to the University of Florida Proton Therapy Institute, where they held multiple positions including Administrative Manager from September 2005 to October 2011 and later became Director of Admin Services and Employee Benefits. Currently, Holly Mostoller serves as the Director of Human Resources at the University of Florida Proton Therapy Institute, a position they have held since July 2014.

Holly Mostoller attended Bowling Green State University from 1990 to 1994, where they studied Biological Science and Environmental Policy. Their degree is unspecified. Additionally, Holly obtained a Professional in Human Resources® (PHR®) certification from the Human Resources Certification Institute in October 2020. Prior to that, in July 2015, they acquired a SHRM - Certified Professional (SHRM-CP) certification from SHRM.