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Gregory Smith has a diverse work experience starting in 1989 as a Research Intern at Amgen. Gregory then went on to work as a Graduate Student Researcher at the University of Pennsylvania from 1990 to 1996. In 1996, they became a Postdoctoral Research Fellow at Princeton University, where they worked until 2001. Following this, they joined Northwestern University - The Feinberg School of Medicine as a Professor in October 2001. Most recently, they have held the position of President at Thyreos since July 2017.

Gregory Smith attended UC Santa Barbara from 1985 to 1990, where they obtained a Bachelor of Arts degree in Microbiology, with a focus on General Microbiology. Following this, they pursued further education at the University of Pennsylvania from 1990 to 1996. During this time, they completed a Doctor of Philosophy (PhD) in Medical Microbiology and Bacteriology. Lastly, from 1996 to 2001, Gregory attended Princeton University, although no specific degree is mentioned. However, their field of study at Princeton University was Virology.