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James R. Downing is the current President & CEO at St. Jude Children's Research Hospital. James has held this position since January 2011. Prior to that, they served as the Deputy Director of the hospital from January 2004 to December 2014. James also served as the Scientific Director and Executive Vice President from January 2004 to December 2014. Prior to that, they were with St. Jude Children's Research Hospital for over 10 years in various positions, including Chair of the Department of Pathology from January 1997 to December 2009, and Associate Member of the Department of Pathology from January 1991 to December 1997. James started their career at St. Jude Children's Research Hospital as an Assistant Member of the Department of Pathology and Laboratory Medicine from January 1986 to December 1991.

James R. Downing has a Doctor of Medicine (M.D.) from the University of Michigan and a Bachelor of Science (B.S.) from the University of Michigan-Dearborn.

Some direct reports include Stephen Gottschalk - Chair, Department of Bone Marrow Transplantation & Cellular Therapy, Carlos Rodriguez-Galindo - EVP & Chair of Department of Global Pediatric Medicine, and Charles W. M. Roberts - EVP & Director of Comprehensive Cancer Center.