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John Coffin holds the post of American Cancer Society Research Professor at the Tufts University School of Medicine and is a special advisor to the director at the National Cancer Institute’s Center for Cancer Research. He serves on a number of national committees to review and set policy regarding retroviruses and disease. John previously served as the founding director of the HIV Drug Resistance Program (renamed as the HIV Dynamics and Replication Program in 2015) in the National Cancer Institute. John was elected to membership in the National Academy of Sciences in 1999. He received his Ph.D. from the University of Wisconsin in the laboratory of Dr. Howard Temin and was a postdoctoral fellow with Dr. Charles Weissmann at the University of Zürich.