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James Ayre has a diverse work experience spanning several prominent financial companies. Their most recent role was as Head of Investments for Charities at Rathbones Group Plc, starting in November 2023. Prior to that, they served as Co-Head of Investments at CCLA Investment Management from May 2007 until November 2023. Before their time at CCLA, James worked at Santander Asset Management as a Senior Investment Analyst from November 2003 to May 2007, and as an Investment Analyst from January 2000 to October 2003. James began their career as a Tax Analyst at Moore Stephens International, where they worked from September 1998 to December 1999.

James Ayre attended the University of Leicester from 1995 to 1998, where they obtained a BA (Hons) degree in History. In 1996-1997, they also attended the Robert Manning School, although specific details about the degree and field of study are not available. Additionally, James Ayre has obtained several certifications, including CFA ESG from the CFA Institute in January 2023, CIMA - Certificate in Business Accounting from CIMA in October 2017, and the CISI Diploma, CISI - Regulation & Compliance, CISI - Fund Management, and CISI - Private Client Investment Advice & Management from The Chartered Institute for Securities & Investment (The CISI) in various years ranging from 2011 to 2012. There is also mention of an Investment Management Certificate (IMC), although no specific details about its acquisition are provided.