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Pursuing several lines of research related to the role of serotonin-signaling in the cortex, demonstrating an important role for cortical influence on behaviors related to schizophrenia and anxiety. His lab also has an active program examining the role of the neurotransmitter serotonin on the development of brain circuits that modulate affective and anxiety states. Additionally, the group has developed a mouse model of epigenetic effects of paternal age on behavior and brain function.