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James McMillan has a solid background in finance and operations management. James started their career as an Assistant Treasurer at the Council for a Parliament of the World's Religions in 1993 and then worked as a Senior Accountant at the Illinois Institute of Technology. In 1998, they became the Finance Director at Sarah's Inn, where they oversaw financial operations for a social service agency. McMillan's next position was Director of Finance at Rainbow House from 2001 to 2003, where they played a crucial role in restructuring the organization's operations. Since 2003, they have been the Chief Financial Officer and Vice President of Operations at Equip for Equality, where they manage all financial affairs and oversees budget development and human resources.

James McMillan's education history includes a Master of Nonprofit Administration degree from North Park University, which they obtained between 2015 and 2018. James also pursued a Master of Business Administration (MBA) degree from the University of St. Francis, although no specific start or end year is provided. Additionally, they participated in the Non-Profit Leadership Program at the Kellogg Business School, although no specific dates are mentioned. James engaged in graduate studies in Non-Profit Management at Saint Xavier University, and for their undergraduate studies, they attained a Bachelor of Business Administration (BBA) degree with a specialization in Accounting from Loyola University Chicago.