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Jonathan Miller started their work experience as a Flight Student for the US Navy in March 2011 and completed this role in May 2014. Jonathan then joined Atlas Coffee Club as a Co-Founder in December 2014. In April 2005, they began working at Purdue University until April 2009. In June 2008, they had a brief internship as a Stress Intern at Hawker Beechcraft. Returning to the US Navy, Jonathan worked as the Squadron Maintenance Division Officer and Assistant Operations Officer from June 2014 to April 2017. Jonathan then became a TOPGUN Instructor from May 2017 to March 2021, during which time they led efforts to standardize fighter tactics and identify F-35 platform capability gaps. Jonathan's latest role is as the F-35 Training Officer for the US Navy, starting in April 2021.

Jonathan Miller attended Purdue University from 2005 to 2009, where they pursued a degree in Engineering with a focus on Aeronautics. Jonathan completed their studies in Aeronautical Engineering and Aerodynamics. In May 2023, Jonathan earned a certification in Virtual Reality and Augmented Reality from Emeritus.